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The antioxidant activity of PQQ under pro-oxidative stressors seems to be also related to other mechanisms, such as the stimulation of many signaling pathways promoting the antioxidant defense and scavenging dangerous ROS, as phosphatidylinositol 3-kinase/Akt, Nrf2, and DJ-1 pathways, or by the modulation of ROS-producing receptors, as N-methyl-D-aspartate (NMDA) receptors (Aizenman et al., 1992, 1994

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